their degree of true altruism ...

True extent of their altruism, and in accordance with this have an impact on the "selfish gene, isolating it from the mutually beneficial exchange of altruistic, or giving him an additional selective pressure. The level of true altruism in such a population may be higher or less high, but it will not be zero (relative to the level of kin altruism), since there would be zero selective pressure on gene selfishness (or otherwise, will not be zero selective advantage of true altruists are able to find each other for mutually beneficial exchange of altruistic). Consequently, contradicted one of the principles of the modern concept of evolution - the maximization of inclusive fitness (in the modern sense of the word-lit. № 5) If the individual is capable of behavioral characteristics to determine with some degree of certainty, the presence of "altruism gene" from other individuals, then relation to their relatives, as possible recipients it will have a "correction" on these signs. This specimen, the donor will donate for the benefit of a relative, as a more likely carrier of the gene of altruism, to a greater extent than in favor of another with the same coefficient of genetic relationship, in favor of one of the brothers is greater than the benefit of another. This individual will donate part of their fitness or fitness of the whole (or, equivalently, for example, fitness of their two relatives with the genetic similarity coefficient of r = 0,5) for the benefit of individuals - recipients, the probability of carriers of the gene "altruism" factor genetic relationships in which this particular donor can be close to zero. According to modern concepts of the theory of evolution, this individual will selectively lose. But the carriers of the gene of altruism - the relatives of altruism gene "will be distributed in the population more rapidly, displacing the non-native speakers. Consider the model of this process in more detail. Sheet 7 Suppose there is a population consisting of genotypes A, B, C. etc. Each of these genotypes has some single genes - we denote them 1,2,3,4. (Ie, genotype A has genes A1, A2, A3, A4, genotype B - B1, B2, B3, B4 and m . etc.). In accordance with the theory of kin-selection, each individual is selectively advantageous to worry not only about the distribution of gene carrier to which it is, but also the identical genes - genes that are available in close and distant relatives. Let's say one of the individuals of genotype A appeared reciprocal altruism gene - a gene A5, a gene inducing him to sacrifice for the benefit of another individual, the bearer of the same gene, excluding the degree of genetic relatedness. Not whether it will selectively lose in that case? Such a reciprocal altruist (which can be called "seeing altruist") will give the benefit of any individual, any genotype (A, B, C, etc.) having the gene A5, and in favor of unknown genes, which can be designated as genes O (ie, it will be donating, the donor, in favor of the recipient, whose genes are O1, O2, O3, O4, A5.). In turn, this recipient is O, having the gene of A5, becoming a donor, will also give the benefit of individuals having a gene of A5 and other unknown genes (ie,