If a victim podrazumevaetsya ...

If a victim podrazumevaetsya any service, assistance from the donor, it is clear that such a service could be for the recipient many times more valuable than the efforts of donor spent on the provision of services (for example, rescue a drowning man). But it may it be beneficial to decrease the viability of the donor recipient? Decrease in the viability of individuals may be beneficial for other individuals just like the opportunity to receive additional resources, BUSINESS donor (food and others). But how can the resources of X, "saved" by lowering the body's ability to live donor, to benefit the recipient more than the X / r (for the brother, sister - more than 2 times, for cousins, more than 8 times etc.) From the standpoint of kin-selection that seems completely impossible. In fact, it is not. Page 2 The fact that the common scheme of kin-selection is to some extent, simplistic. It notes, for example, Dawkins (lit. № 3), proposing to introduce an amendment to the "life expectancy" or "general ability to favor their own genes in the course of their lives." Indeed, although in relation to his grandfather lying on the stove, it viripotent grandson, running around the street, has a coefficient of genetic relatedness r, equal to 1 / 4, but the effectiveness of the spread of their own genes (some of which they have identical) they will be completely different. For my grandfather saved his own life at the cost of the lives of three (and more than four) grandchildren will most likely not selectively advantageous thing to do. To more accurately reflect the selective advantage obtained altruistic donor-selection process, you must apply the rate of conversion efficiency of the resource in a special fitness. Under the resource should include not only food resources, but all that may increase the fitness of individuals - better, more comfortable, safe and t.p.Peredacha any resource such as a piece of meat, the brother would not be selectively advantageous thing to do. But it will not, and a complete loss for the donor. Since the coefficient r between them is 1 / 2, with a selective point of view can be considered such a thing as a loss of 50% of the resource. But the conversion efficiency of the resource in the fitness of the brothers may be quite different, for example, due to the varying efficiency of digestive system, or for other reasons. The coefficient K should show the degree of selective advantage, which is able to get the specimen due to a unit of resource consumption. This ratio should reflect not a temporary, short-term losses (or increase) the conversion efficiency of the resource in a device which can be, for example in the case of relatively short-term illness, and the average conversion efficiency of the resource in the inclusive fitness for all life, from the time frame (from the possible altruistic Act). The coefficient K should show on what value can be enhanced reproductive value of individuals of age (age-specific reproductive value - lit. № 5) by receiving a unit of resource (of course taking into account the contribution to inclusive fitness).